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What is the function of the substitutes in the doctoral committee?

The substitutes of your doctoral committee (one substitute for the university lecturers and one for the academic staff) will stand in if a member of your doctoral committee is unable to attend at very short notice (e.g. due to illness).

Since your defense can only take place with a complete doctoral committee (4 professors and 1 academic staff member), substitutes should always be named by the day of the defense.