AG Hilger (Morphologie und Systematik) - Publikationen

An dieser Stelle präsentieren wir Ihnen Publikationen der AG Morphologie und Systematik der Phanerogamen unter der Leitung von Prof. Dr. Hartmut H. Hilger.


Unser Hauptarbeitsgebiet: Boraginaceae s.l. ab 2000.

Alle weiteren Publikationen sind in den entsprechenden Jahresberichten aufgeführt (ab 1988).


Publikationen zu weitergefassten Themenkomplexen und Kooperationen

Monographien, Floren, Bücher, ...



Poster der Arbeitsgruppe, auf denen Forschungsergebnisse im Rahmen von Tagungen und Kongressen vorgestellt wurden.


Preise und Auszeichnungen

Katharina-Heinroth-Preise der Gesellschaft naturforschender Freunde Berlin für Diplom-Arbeiten über Boraginales aus der AG

2001: Marc Gottschling (2000): Phylogenetische Analyse einiger Vertreter der Ehretiaceae Mart. ex Lindl. (Boraginales).

2005: Natalie Cusimano (2004): Untersuchungen an Onosma (Boraginaceae) von Serpentinstandorten in der Umgebung von Florenz (Italien).

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Publikationen ab dem Jahre 2000

Boraginales (Boraginaceae s.l.)

Federico Luebert, Lorenzo Cecchi, Michael W. Frohlich, Marc Gottschling, C. Matt Guilliams, Kristen E. Hasenstab-Lehman, Hartmut H. Hilger, James S. Miller, Moritz Mittelbach, Mare Nazaire, Massimo Nepi, Daniele Nocentini, Dietrich Ober, Richard G. Olmstead, Federico Selvi, Michael G. Simpson, Karel Sutorý, Benito Valdés, Genevieve K. Walden & Maximilian Weigend (2016): Familial classification of the Boraginales. - Taxon 65(3):

Weigend, M., Luebert, F., Gottschling, M., Couvreur, T.L.P., Hilger, H.H. & Miller, J.S. (2014): From capsules to nutlets — phylogenetic relationships in the Boraginales. - Cladistics 30(5): 508-518. DOI: 10.1111/cla.12061

Hilger, H.H., Selvi, F., Bigazzi, M., Langstrom, E., Gottschling, M., Zippel, E., Diane, N. & Weigend, M. (2005): The Euro+Med treatment of Boraginaceae - a response. - Willdenowia 35 (1): 43-48.

Gottschling, M., Diane, Nadja, Hilger, H.H. & Weigend, M. (2004): Testing hypotheses on disjunctions present in the Primarily Woody Boraginales: Ehretiaceae, Cordiaceae, and Heliotropiaceae, inferred from ITS1 sequence data. - Int. J. Plant Sci. 165 (4 Suppl.): S123-S135.

Hilger, H.H. & Förther, H. (2004): „Boraginaceae“ - in: Neotropical Plant Families - ed. New York Botanical Garden.

Buys, M.H. & Hilger, H.H. (2003): Boraginaceae cymes are exclusively scorpioid and not helicoid. - Taxon 52: 719-724.

Diane, Nadja, Hilger, H.H. & Gottschling, M. (2002): Transfer cells in the seeds of Boraginales . - Bot. J. Linn. Soc. 140: 155-164.

Gottschling, M., Hilger, H.H., Wolf, M. & Diane, Nadja (2001): Secondary Structure of the ITS1 Transcript and its Application in a Reconstruction of the Phylogeny of Boraginales. - Plant biol. 3: 629-636

Hilger, H.H. & Zippel, Elke (2001): "Studies in the Boraginaceae" - An Index to the Publications of Ivan M. JOHNSTON dealing with the Borage Family. - Haussknechtia Beiheft 11: 1-151. Jena

Hofmann, Maria (2004): "Hydrophyllaceae" - in: Neotropical Plant Families - ed. New York Botanical Garden.

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Jeiter, J.; Danisch, F.& Hilger, H.H. (2016): Polymery and nectary chambers in Codon (Codonaceae) - flower and fruit development in a small, capsule-bearing family of Boraginales. - Flora 220: 94-102.

Weigend, M. & H.H. Hilger (2010): Codonaceae — a newly required family name in Boraginales. - Phytotaxa 10: 26-30.

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Cordiaceae und Ehretiaceae

Gottschling, M., Nagelmüller, S. & Hilger, H.H. (2014): Generative ontogeny in Tiquilia (Ehretiaceae, Boraginales) and phylogenetic implications. - Biol. J. Linn. Soc. 112: 520-534

Gottschling, M., F. Luebert, H.H. Hilger & J. Miller (2014): Molecular delimitations in the Ehretiaceae (Boraginales). - Mol. Phylogen. Evol. 72: 1-6.

Gottschling, M. & Miller, J.S. (2007): A revision of Bourreria (Ehretiaceae, Boraginales) in South America. - Ann. Mo. Bot. Gard. 94 (4): 734-744.

Gottschling, M. & Miller, J.S. (2007): Typification of Bourreria names (Ehretiaceae, Boraginales) based on specimens collected by Charles Wright in Cuba. - Taxon 56 (1): 237-242.

Miller, J.S. & Gottschling, M. (2007): Generic classification in the Cordiaceae (Boraginales): resurrection of the genus Varronia P.Br.- Taxon 56 (1): 163-169.

Gottschling, M. & Miller, J.S. (2006): Clarification of the taxonomic position of Auxemma, Patagonula, and Saccellium (Cordiaceae, Boraginales). - Syst. Bot. 31 (2): 361-367.

Gottschling, M. & Miller, J.S. (2005): A new species of Bourreria (Ehretiaceae, Boraginales) from Costa Rica. - Novon 15: 425-428.

Gottschling, M., Miller, J.S., Weigend, M. & Hilger, H.H. (2005): Congruence of a phylogeny of Cordiaceae (Boraginales) inferred from ITS1 sequence data with morphology, ecology, and biogeography. - Ann. Missouri Bot. Gard. 92 (3): 425-437.

Gottschling, M. (2004): Floral ontogeny in Bourreria (Ehretiaceae, Boraginales). - Flora 199 (5): 409-423.

Gottschling, M. & Hilger, H.H. (2004): Characterization of a novel fruit type found in Ehretia (Ehretiaceae, Boraginales). - Blumea 49: 145-153.

Gottschling, M. & Hilger, H.H. (2004): The systematic position of Ehretia cortesia nom. nov. (syn. Cortesia cuneifolia: Ehretiaceae, Boraginales) inferred from molecular and morphological data. - Taxon 53 (4): 919-923.

Gottschling, M. & Hilger, H.H.(2003): First fossil record of transfer cells in angiosperms. - Amer. J. Bot. 90 (6): 957-959.

Gottschling, M., Mai, D.H. & Hilger, H.H. (2002): The systematic position of Ehretia fossils (Ehretiaceae, Boraginales) from the European Tertiary and implications for character evolution. - Review of Palaeobotany and Palynology 121 (2): 149-156.

Gottschling, M. & Hilger, H.H. (2001): Phylogenetic analysis and character evolution of Ehretia and Bourreria (Ehretiaceae, Boraginales) and their allies based on ITS1 sequences. - Bot. Jahrb. Syst. 123: 249-268.

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Luebert, F. & H.H. Hilger (2014): Typification of Heliotropium and Tournefortia (Heliotropiaceae) species described by Ruiz and Pavón. - AJB Madrid 71 (2) epub doi: http://dx.doi.org/10.3989/ajbm.2332.
Luebert, F.; Jacobs, P.; Hilger, H.H. & Muller, L.A. (2013 online): Evidence for non-allopatric speciation among closely related sympatric Heliotropium species in the Atacama Desert. - Ecol. Evol. online: http://onlinelibrary.wiley.com/doi/10.1002/ece3.929/full.

Luebert, F., G. Brokamp, J. Wen, M. Weigend & H.H. Hilger (2011): Phylogenetic relationships and morphological diversity in Neotropical Heliotropium (Heliotropiaceae). - Taxon 60: 663-680.

Luebert, F., H.H. Hilger & M. Weigend (2011): Diversification in the Andes: Age and origins of South American Heliotropium lineages (Heliotropiaceae, Boraginales. - Mol. Phylogenet. Evol. 61: 90-102.

Luebert, F, M Weigend & H H Hilger (2010): Epitypification of Heliotropium arborescens L. (Heliotropiaceae). - Taxon 59: 1263-1266.

Luebert, F. & Wen, J. (2008): Phylogenetic analysis and evolutionary diversification of Heliotropium Sect. Cochranea (Heliotropiaceae) in the Atacama Desert. - Syst. Bot. 33 (2): 390-402.

Diane, Nadja, Jakob, Claudia & Hilger, H.H. (2003): Comparative leaf anatomy and distribution of foliar trichome types in Heliotropiaceae - Flora 198: 468-485.

Hilger, H.H. & Diane, Nadja (2003): A systematic analysis of Heliotropiaceae (Boraginales) based on trnL and ITS1 sequence data. - Bot. Jahrb. Syst. 125 (1): 19-51.

Weigend, M., Brokamp, G., Kirbach, A. & Förther, H.(2003): Notas sobre Heliotropium krauseanum Fedde, la unica especie de Heliotropium Sect. Cochranea del Perú (Heliotropiaceae = Boraginaceae subfam. Heliotropioideae). - Arnaldoa 10 (1): 61-74.

Diane, Nadja, Förther, H. & Hilger, H.H. (2002): Molecular Systematics of the Heliotropiaceae (Boraginales) Inferred from ITS1 Sequences. - Amer. J. Bot. 89 (2): 287- 297.

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Hydrophyllaceae incl. Namaceae

Hofmann, Maria, Walden, Genevieve K., Hilger, H.H. & Weigend, M. (2016): Hydrophyllaceae. pp. 221-238 in Kadereit, J.W. & Bittrich, V. (eds.): The Families and Genera of Vascular Plants Vol. 14: Flowering Plants. Eudicots. Aquifoliales, Boraginales, Bruniales, Dipsacales, Escalloniales, Garryales, Paracryphiales, Solanales (except Convolvulaceae), Icacinaceae, Metteniusaceae, Vahliaceae. Springer - Switzerland

Boraginaceae (s.str.)

Chacon, J., Luebert, F., Hilger, H.H., Ovchinnikova, Svetlana, Selvi, F., Cecchi, L., Guilliams, C.M., Hasenstab-Lehman, Kristen, Sutorý, K., Simpson, M.G., Weigend, M. (2016): The borage family (Boraginaceae s.str.): A revised infrafamilial classification based on new phylogenetic evidence, with emphasis on the placement of some enigmatic genera. - Taxon 65 (3): 523-546.
Weigend, M., Luebert, F., Selvi, F., Brokamp, G. & H.H. Hilger (2013): Multiple origins for Hounds tongues (Cynoglossum L.) and Navel seeds (Omphalodes Mill.) – the phylogeny of the borage family (Boraginaceae s.str.). - Molecular Phylogenetics and Evolution 68(3): 604-618.

Muller, L.A.H. & H.H. Hilger (2015): Insights into the effects of serpentine soil conditions on the community composition of fungal symbionts in the roots of Onosma echioides. - Soil Biol. Biochem. 81: http://www.sciencedirect.com/science/article/pii/S0038071714003745
Cecchi, L., A. Coppi, H.H. Hilger & F. Selvi (2014): Non-monophyly of Buglossoides (Boraginaceae: Lithospermeae): Phylogenetic and morphological evidence for the expansion of Glandora and reappraisal of Aegonychon. - Taxon 63: 1965-1078

Weigend, M., M. Gottschling, F. Selvi & H.H. Hilger (2009): Marbleseeds are gromwells - Systematics and evolution of Lithospermum and allies (Boraginaceae tribe Lithospermeae) based on molecular and morphological data. - Mol. Phylogenet. Evol. 52: 755-768.

Weigend, M., Gotrschling, M., Hilger, H.H. & Nürk, N.M. (2010): Five new species of Lithospermum L. (Boraginaceae tribe Lithospermeae) in Andean South America . another radiation in the Amotape-Huancabamba Zone. - Taxon 59: 1161-1179.

Pustovoytov, K., Riehl, S., Hilger, H.H. & Schumacher, E. (2010): Oxygen isotopic composition of fruit carbonate in Lithospermeae and its potential for paleoclimate research in the Mediterranean. - Global Planet. Change 71: 258-268.

Weigend, M., Gottschling, M., Selvi, F. & Hilger, H.H. (2009): Marbleseeds are gromwells - Systematics and evolution of Lithospermum  and allies (Boraginaceae tribe Lithospermeae) based on molecular and morphological data. - Molec. Phylogen. Evol. 52 (3): 755-768.

Thomas, D.C., Weigend, M. & Hilger, H.H. (2008) Phylogeny and systematics of Lithodora (Boraginaceae, Lithospermeae) and its affinities to the monotypic genera Neatostema, Mairetis, Halacsya and Paramoltkia based on ITS1 and trnLUAA-sequence data and morphology. - Taxon 57 (1): 79-97.

Mengoni, A., Selvi, F., Cusimano, N., Galardi, F., Gonnelli, C. (2006): Genetic diversity inferred from AFLP fingerprinting in populations of Onosma echioides (Boraginaceae) from serpentine and calcareous soils. - Plant Biosystems 140 (2): 211-219.

Clermont, Anja, Zippel, Elke & Hilger, H.H. (2003): Verbreitung und Differenzierung der mitteleuropäischen Unterarten von Buglossoides arvensis (L.) I.M.Johnst. (Boraginaceae). - Feddes Repert. 114 (1-2): 58-70.

Hilger, H.H. & Böhle, Uta-Regina (2000): Pontechium - a new genus separated from Echium and Lobostemon (Boraginaceae). - Taxon 49: 737-746.

Böhle, Uta-Regina, Hilger, H.H. & Martin, W. (1996): Island colonization and evolution of the insular woody habit in Echium L. (Boraginaceae). - Proc. Natl. Acad. Sci. USA 93: 11740-11745.

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Weigend, M., M. Gottschling, F. Selvi & H.H. Hilger (2010): Fossil and extant Western Hemisphere Boragineae, and the polyphyly of Trigonotideae Riedl (Boraginaceae: Boraginoideae). - Syst. Bot. 35: 409-419.

Selvi, F., Bigazzi, M., Hilger, H.H., & Papini, A. (2006): Molecular phylogeny, morphology and taxonomic re-circumscription of the generic complex Nonea/Elizaldia/Pulmonaria/Paraskevia (Boraginaceae - Boragineae). - Taxon 55 (4): 907-918.

Bigazzi, M., Selvi, F. & Hilger, H.H. (2004): Nonea pisidica (Boraginaceae - Boragineae), a new species from southw est Anatolia and its relationships inferred from karyology and cpDNA sequences. - Plant Biosystems 138 (2): 135-144.

Hilger, H.H., Selvi, F., Papini, A. & Bigazzi, M. (2004): Molecular systematics of Boraginaceae tribe Boragineae based on ITS1 and trnL sequences, with special reference to Anchusa s.l. - Ann. Bot. 94 (2): 201-212.

Selvi, F., Papini, A., Hilger, H.H., Bigazzi, M. & Nardi, E. (2004): The phylogenetic relationships of Cynoglottis (Boraginaceae - Boragineae) inferred from ITS, 5.8S and trnL sequence variation - Pl. Syst. Evol. 246 (3-4): 195-209.

Bigazzi, M., Hilger, H,H. & Selvi, F. (2002): Evidence from nuclear and chloroplast DNA for the placement of Anchusa macedonica in the genus Gastrocotyle (Boraginaceae). - Webbia 57 (2): 173-180.

Danin, A. (2001): Hormuzakia negenvensis(Danin) Danin & Hilger comb. nov., pp. 110-111 in Danin, A.: The nomenclature news of Flora Palaestina. - Fl. Medit. 10: 109-172.

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Holstein, N., Hilger, H.H. & Weigend, M. (in press, 2016 accepted): "Omphalodes" (Boraginaceae) in the South America and the South Pacific". - Phytotaxa

Hilger, H.H., Greuter, W. & Stier, V. (2015): Taxa and names in Cynoglossum sensu lato (Boraginaceae, Cynoglosseae): an annotated, synonymic inventory, with links to the protologues and mention of original material. - Biodiversity Data Journal 3: e4831 (22 Apr 2015) doi: 10.3897/BDJ.3.e4831.
König, J., Siwakoti, M., Hilger, H.H. & Weigend, M. (2015): A revision of the genus Cynoglossum L. (Boraginaceae Juss.) in Nepal and notes on the widespread Asian species. - Phytotaxa 224(1): 1-28. http://biotaxa.org/Phytotaxa/article/view/phytotaxa.224.1.1/18191
Dickoré, W.B. & Hilger, H.H. (2015): Decalepidanthus (Boraginaceae) includes and antedates Pseudomertensia; a synopsis of the genus. - Phytotaxa 226(2): 131-143.

Greuter, W., Hilger, H.H. & Stier, V. (in press): Proposal to conserve the name Omphalodes verna against Omphalodes omphlaoides (Boraginaceae). - Taxon.

Hilger, H.H., Cole, T.C.H. & Selvi, F. (2014): J.G.C. Lehmann's 'Botanical Observations' of 1818 on Coldenia, Colsmannia, Cynoglossum, and Omphalodes (commented translation). Biodiversity Data Journal 2: e1064.

Weigend, M., Luebert, F., Selvi, F., Brokamp, G. & Hilger, H.H. (2013): Multiple origins for Hound’s tongues (Cynoglossum L.) and Navel seeds (Omphalodes Mill.) – The phylogeny of the borage family (Boraginaceae s.str.). - Mol. Phylogen. Evol. 68: 604-618.

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The Last Millenium

Hofmann, Maria (1999): Flower and fruit development in the genus Phacelia (Phacelieae, Hydrophyllaceae): characters of systematic value. - Syst. Geogr. Pl. 68(1-2): 203-212

Böhle, Uta-Regina, Hilger, H.H., Cerff, R. & Martin, W. (1994): Non-coding chloroplast DNA for plant molecular systematics at the infrageneric level - In: Schierwater, B., Streit, B., Wagner, G.P. & DeSalle, R. (eds.): Molecular Ecology and Evolution: Approaches and Application: 391- 403.

Hilger, H.H. (1994): Die Evolution der Rauhblattgewächse (Boraginaceae) in Makaronesien. - Ulmensien 8: 23- 32.

Hilger, H.H., Hoppe, J.R. & Hofmann, Maria (1993): Energiedispersive Röntgenmikroanalyse (EDX) von Boraginaceae subfam. Boraginoideae - Klausenoberflächen. Sind Silicium- und Calcium- Einlagerungen in die Fruchtwand systematisch verwertbare Merkmale? - Flora 188: 387- 398.

Hilger, H.H. (1992): Morphology of Heliotropium (Boraginaceae) dispersal units. - Bot. Acta 105: 387- 393.

Hilger, H.H. (1989): Flower and fruit development in the Macaronesian endemic Ceballosia fruticosa (syn. Messerschmidia fruticosa, Boraginaceae, Heliotropioideae). - Pl. Syst. Evol. 166: 119 - 129.

Briechle, Monika & Hilger, H.H. (1988): Die Embryogenese von Microparacaryum intermedium und die embryologische Klassifizierung der Boraginaceae nach Souèges - Flora 181: 45 - 59.

Hilger, H.H. (1987): Flower and fruit development in Wigandia caracasana (Hydrophyllaceae). - Amer. J. Bot. 74: 250 - 259.

Hilger, H.H. (1987): Fruchtbiologische Untersuchungen an Heliotropioideae (Boraginaceae). I. Die Ontogenie der monospermen Früchte von Heliotropium supinum L. - Flora 179: 291 - 303.

Hilger, H.H. (1986): Mikromorphologische Untersuchungen an Klausenoberflächen von Boraginaceae II. Mattiastrum (Boiss.) Brand (Cynoglosseae). - Beitr. Biol. Pflanzen 61: 145 - 165.

Hilger, H.H. (1985): Ontogenie, Morphologie und systematische Bedeutung geflügelter und glochidientragender Cynoglosseae- und Eritrichieae-Früchte (Boraginaceae). - Bot. Jahrb. Syst. 105: 323 - 378.

Hilger, H.H., Balzer, Mechtild, Frey, W. & Podlech, D. (1985): Heteromerikarpie und Fruchtpolymorphismus bei Microparacaryum, gen. nov. (Boraginaceae). - Pl. Syst. Evol. 148: 291 - 312.

Hilger, H.H. (1984): Wachstum und Ausbildungsformen des Gynoeceums von Rochelia (Boraginaceae). - Pl. Syst. Evol. 146: 123 - 139.

Hilger, H.H. & Richter, Ursula (1982): Untersuchungen zur Ausbildung der Klausen und ihrer Oberflächenskulpturen bei Paracaryum intermedium (Boraginaceae) aus Jordanien. - Beitr. Biol. Pflanzen 57: 205 - 220.

Hilger, H.H. (1981): Die Perikarpentwicklung geflügelter Klausen aus dem Paracaryum - Verwandtschaftskreis (Cynoglosseae, Boraginaceae). - Pl. Syst. Evol. 138: 73 - 88.

Hilger, H.H. (1981): Entwicklungstendenzen im Fruchtbau der anemochoren Boraginaceengattungen Mattiastrum und Paracaryum. - Ber. Deutsch. Bot. Ges. 94: 677 - 687.